In New York, the simple and naive just-graduated in journalism Andrea Sachs is hired to work as the second assistant of the powerful and sophisticated Miranda Priestly, the ruthless and merciless executive of the Runway fashion magazine. Andrea dreams to become a journalist and faces the opportunity as a temporary professional challenge. The first assistant Emily advises Andrea about the behavior and preferences of their cruel boss, and the stylist Nigel helps Andrea to dress more adequately for the environment. Andrea changes her attitude and behavior, affecting her private life and the relationship with her boyfriend Nate, her family and friends. In the end, Andrea learns that life is made of choices.
The story tells the professional adventure of Andrea, whose greatest dream is to become a journalist. Andrea gets a job in the fashion industry through Runway magazine, the most famous of its type, to make ends meet. But Andrea won't develop her writing skills in the magazine, but her talents as the editor in chief's assistant, Miranda. The problem is that Miranda is a merciless, posh and cruel woman, making the experience a living hell for the girl. The environment in the place will be cold and extremely critical with the physical appearance. The girl will have to change her simple and plain style, for a more trendy and elegant one, in order to gain the acceptance of her ruthless boss and colleagues, specially Emily, her unpleasant workmate. Despite everything against Andrea in the office, she will consider the experience as a challenge, drastically changing her clothes and self-image, with the help of Nigel, the magazine's art director. Nevertheless, the job becomes extremely demanding, because of Miranda's tough work rhythm and nearly impossible tasks, leaving Andrea without a private life with her boyfriend, family and friends. Maybe the old Andrea has gone, now more preoccupied about her image and her future in the magazine.
Miranda Priestley, of "Runway" magazine tears up the landscape as a demanding fashion editor. She is a terror to everyone who is around her as is quickly depicted in the opening scenes of the movie. Her first assistant strives to please her and tries to emulate her, but one can sense that she is not quite as hard as she tries to put on. Into this mix comes a young woman who knows nothing of the fashion industry, has never read the magazine, and doesn't know who Miranda Priestley is. She only sees this as a stepping stone to another journalism position. Showing no fashion sense and immediately scorned by everyone, Miranda nonetheless hires her as the second assistant. When Miranda demands that she obtain the next unpublished Harry Potter manuscript, you can sense that she is trying to force her to quit, but it makes the young woman dig in to please her boss. With the help of one of the magazine's fashion editors, she gets a complete makeover and a new security. However, with her new appearance and the demands placed on her, she starts to lose her friends, family and her live-in boy friend. As she is whisked away to Paris with Miranda and faces all of the glamor that could be hers, including a flashy if not artificial freelance journalist, she is forced to make the decision of where she wants to be in her life
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